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Courteney Cox nació y se crio en Birmingham, Alabama, un suburbio de Mountain Brook1 ,2 hija del empresario Richard Lewis Cox (28 de enero, 1931 – 3 de septiembre, 2001) y de su esposa Courteney (nacida bajo Bass, luego, Copeland). Tiene dos hermanas mayores, Virginia y Dottie, y un hermano mayor, Richard, Jr. Sus padres se divorciaron en 1974 y su madre se casó de nuevo con un empresario llamado Hunter Copeland (tío de Ian Copeland). Después de graduarse de la secundaria Mountain Brook, Cox se fue para la universidad Mount Vernon College en Washington, D.C. pero no completó su curso de arquitectura, optando en su lugar a seguir una carrera de modelaje y actuación.

 

Cox estudió arquitectura en la universidad Mount Vernon para Mujeres en Washington, D.C., que ahora es parte de la universidad Georg Washington, antes de que se fuera. Salió por primera vez en prominencia en un vídeo musical para Bruce Springsteen, “Dancing in the Dark” (en que ella era la chica que fue sacada al escenario para bailar con Springsteen). Cox es notable por ser la primera persona en usar la palabra “período” en la televisión de Estados Unidos para referirse a la menstruación en una campaña publicitario de 1985 para los tampones Tampax. Sus primeros papeles en el cine incluyen Masters of the Universe (1987) y Cocoon: The Return (1988). Sus primeros trabajos en televisión incluyen un papel protagonista en la serie Misfits of Science (1985), y luego un papel recurrente (1987-89) en la serie Family Ties como Lauren Miller, la novia de Alex P. Keaton (Michael J. Fox). Tuvo un papel secundario como Jewel, una asistente de uñas para el personaje de James Belushi, en la película de fantasía de 1990 Mr. Destiny. En 1994, luego de su debut de la comedia Friends, Cox apareció con Jim Carrey en la película Ace Ventura: Pet Detective y en Seinfeld, en el episodio “The Wife” como la novia de Jerry llamada Meryl.

Más tarde, en 1994, a Cox se le preguntó para una audición para el papel de Monica Geller en una nueva serie de comedia llamada Friends. Al principio el miembro más famoso del nuevo show, Cox se unió con sus compañeros de reparto Jennifer Aniston (Rachel Green), Lisa Kudrow (Phoebe Buffay), Matt LeBlanc (Joey Tribbiani), Matthew Perry (Chandler Bing) y David Schwimmer (Ross Geller) para lo que se convertiría en su papel más famoso, durando en 10 temporadas hasta el 2004. De acuerdo con el Libro Guinness (2005), Cox (junto con sus coprotagonistas femeninas) se convirtieron en las actrices mejor pagadas de la televisión de todos los tiempos cobrando 1 millón de dólares por episodio, durante las dos últimas temporadas de Friends.

Entre las temporadas quinta y sexta, Cox se casó con David Arquette y como consecuencia cambió su apellido por el de Arquette. En una referencia en broma a ésto se hizo en los créditos de apertura del episodio “The One After Vegas”, dónde el resto del elenco y los productores ejecutivos tenían “Arquette” agregado a sus nombres. La dedicación “Para Courteney y David, que sí se casaron” -una referencia a la decisión de Monica y Chandler de no casarse en el episodio- aparece durante el final del episodio. Una referencia en broma de su segundo nombre “Cox”, fue usado en el episodio de Scrubs, “My Saving Grace”, en que el personaje Taylor Maddox describe al nombre de Cox como “ridículo.”

Después del papel en Friends, Cox fue la primera opción del productor Marc Cherry para un papel estelar como Susan Mayer en Desperate Housewives. Sin embargo, Cox no estaba disponible debido a su embarazo y el papel fue dado a Teri Hatcher. Un par de años después, Cox firmó un contrato con ABC Studios para protagonizar en su propia serie. Desde Friends, Cox se ha concentrado en su familia pero protagonizó en la película independiente November (2005) que tuvo un lanzamiento limitado en los cines, con Tim Allen en Zoom, y en la nueva versión de The Longest Yard como la novia del personaje de Adam Sandler. Ella ofreció su voz para la película animada Barnyard. Se rumoreó de una película para Friends en producción; seguido del éxito de Sex and the City: The Movie, pero ésto fue negado por Warner Bros. y otros.

En 2007, Cox interpretó el papel de Lucy Spiller, una editora de prensa, en Dirt, un drama de televisión por FX. Cox y su esposo David Arquette fueron los productores ejecutivos de la serie. De acuerdo a Cox, la serie fue cancelada después de una segunda temporada en 2008. En julio de 2008, Entertainment Weekly anunció que Cox había sido firmada para estar en tres episodios para la serie Scrubs. En su tercer episodio, ella le dijo al Dr. Cox que Cox era “un nombre ridículo”, en referencia al suyo. En agosto de 2008, se anunció que habría una cuarta película para la serie de Scream, y que querían a los actores originales para representar sus papeles. El 30 de octubre de 2008, TV Guide reportó que Cox estaría protagonizando un piloto para su nueva comedia en ABC llamada Cougar Town de Bill Lawrence. Cox interpreta a una soltera de 40 años de edad, madre, y en busca de nuevas experiencias. Sus co-estrellas son Busy Philipps y Christa Miller. Se reúne con Scott Foley, 10 años después que hicieron Scream 3 juntos. Cox grabó el piloto el 19 de marzo de 2009.19 En junio de 2009, el director de Scream Wes Craven confirmó que ella y Arquette estarían en Scream 4.

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